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Welcome to CIS-BP, the online library of transcription factors and their DNA binding motifs.
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By Model Organism	Arabidopsis_thaliana Caenorhabditis_elegans Danio_rerio Drosophila_melanogaster Homo_sapiens Mus_musculus Neurospora_crassa Rattus_norvegicus Saccharomyces_cerevisiae Xenopus_tropicalis
By Any Species	Acanthamoeba_castellanii Acanthamoeba_polyphaga_mimivirus Acanthocheilonema_viteae Acipenser_baerii Acremonium_chrysogenum Acropora_formosa Acropora_millepora Acyrthosiphon_pisum Aedes_aegypti Aegilops_tauschii Agaricus_bisporus Ailuropoda_melanoleuca Albugo_laibachii Alligator_sinensis Allomyces_macrogynus Alternaria_brassicicola Amanita_muscaria Amborella_trichopoda Amphimedon_queenslandica Anas_platyrhynchos Ancylostoma_caninum Ancylostoma_ceylanicum Ancylostoma_duodenale Angiostrongylus_cantonensis Angiostrongylus_costaricensis Anisakis_simplex Anncaliia_algerae Anolis_carolinensis Anopheles_albimanus Anopheles_arabiensis Anopheles_atroparvus Anopheles_christyi Anopheles_coluzzii Anopheles_culicifacies Anopheles_darlingi Anopheles_dirus Anopheles_epiroticus Anopheles_farauti Anopheles_funestus Anopheles_gambiae Anopheles_maculatus Anopheles_melas Anopheles_merus Anopheles_minimus Anopheles_quadriannulatus Anopheles_sinensis Anopheles_stephensi Antirrhinum_majus Apis_mellifera Aplysia_californica Aptenodytes_forsteri Aquilegia_coerulea Arabidopsis_lyrata Arabidopsis_thaliana Artemia_franciscana Arthrobotrys_oligospora Arthroderma_benhamiae Arthroderma_otae Ascaris_lumbricoides Ascaris_suum Ashbya_gossypii Aspergillus_carbonarius Aspergillus_clavatus Aspergillus_flavus Aspergillus_fumigatus Aspergillus_nidulans Aspergillus_niger Aspergillus_oryzae Aspergillus_ruber Aspergillus_terreus Astyanax_mexicanus Atta_cephalotes Aureobasidium_melanogenum Aureobasidium_pullulans Aureobasidium_subglaciale Aureococcus_anophagefferens Avian_erythroblastosis_virus Avian_musculoaponeurotic_fibrosarco Avian_myeloblastosis_virus Avian_sarcoma_virus_17 Babesia_bovis Batrachochytrium_dendrobatidis Baudoinia_compniacensis Beauveria_bassiana Bigelowiella_natans Biomphalaria_glabrata Bipolaris_maydis Bipolaris_oryzae Bipolaris_sorokiniana Bipolaris_victoriae Bipolaris_zeicola Blastomyces_dermatitidis Blumeria_graminis Boechera_stricta Bombyx_mori Bos_grunniens Bos_taurus Botryobasidium_botryosum Botrytis_cinerea Bovine_papillomavirus_type_2 Brachypodium_distachyon Branchiostoma_floridae Brassica_napus Brassica_oleracea Brassica_rapa Brettanomyces_bruxellensis Brugia_malayi Brugia_pahangi Brugia_timori Buceros_rhinoceros Bursaphelenchus_xylophilus Byssochlamys_spectabilis Caenorhabditis_brenneri Caenorhabditis_briggsae Caenorhabditis_elegans Caenorhabditis_japonica Caenorhabditis_remanei Callithrix_jacchus Calypte_anna Camponotus_floridanus Candida_albicans Candida_dubliniensis Candida_glabrata Candida_guilliermondii Candida_lusitaniae Candida_maltosa Candida_orthopsilosis Candida_parapsilosis Candida_tenuis Candida_tropicalis Canis_familiaris Cannabis_sativa Capitella_teleta Capronia_coronata Capronia_epimyces Capronia_semiimmersa Capsaspora_owczarzaki Capsella_grandiflora Capsella_rubella Carica_papaya Cavia_porcellus Ceriporiopsis_subvermispora Chaetomium_globosum Chaetomium_thermophilum Chelonia_mydas Chlamydomonas_reinhardtii Chlorella_NC64A Chlorella_vulgaris Chlorocebus_sabaeus Choloepus_hoffmanni Chroomonas_mesostigmatica Cicer_arietinum Cimex_lectularius Ciona_intestinalis Ciona_savignyi Citrullus_lanatus Citrus_clementina Citrus_sinensis Cladophialophora_bantiana Cladophialophora_carrionii Cladophialophora_immunda Cladophialophora_psammophila Cladophialophora_yegresii Claviceps_purpurea Clavispora_lusitaniae Clonorchis_sinensis Coccidioides_immitis Coccidioides_posadasii Coccomyxa_subellipsoidea_C169 Cochliobolus_heterostrophus_C5 Colletotrichum_fioriniae Colletotrichum_gloeosporioides Colletotrichum_graminicola Colletotrichum_higginsianum Colletotrichum_orbiculare Colletotrichum_sublineola Coniosporium_apollinis Coprinellus_disseminatus Coprinopsis_cinerea Coprinopsis_scobicola Cordyceps_militaris Crassostrea_gigas Cryphonectria_parasitica Cryptococcus_gattii Cryptococcus_neoformans Cryptosporidium_hominis Cryptosporidium_muris Cryptosporidium_parvum Cucumis_sativus Culex_pipiens Culex_quinquefasciatus Cupiennius_salei Cyanidioschyzon_merolae Cylicostephanus_goldi Cyphellophora_europaea Dacryopinax_sp Dactylellina_haptotyla Danaus_plexippus Danio_rerio Daphnia_pulex Dasypus_novemcinctus Debaryomyces_hansenii Dendroctonus_ponderosae Dichomitus_squalens Dictyocaulus_viviparus Dictyostelium_discoideum Dictyostelium_purpureum Diphyllobothrium_latum Dipodomys_ordii Dirofilaria_immitis Discocelis_tigrina Dothistroma_septosporum Dracunculus_medinensis Drechslerella_stenobrocha Drosophila_ananassae Drosophila_erecta Drosophila_grimshawi Drosophila_melanogaster Drosophila_mojavensis Drosophila_persimilis Drosophila_pseudoobscura Drosophila_sechellia Drosophila_simulans Drosophila_virilis Drosophila_willistoni Drosophila_yakuba Echinococcus_canadensis Echinococcus_granulosus Echinococcus_multilocularis Echinops_telfairi Echinostoma_caproni Edhazardia_aedis Eimeria_tenella Elaeophora_elaphi Eleutheria_dichotoma Emiliania_huxleyi Encephalitozoon_cuniculi Encephalitozoon_hellem Encephalitozoon_intestinalis Encephalitozoon_romaleae Endocarpon_pusillum Entamoeba_dispar Entamoeba_histolytica Entamoeba_invadens Enterobius_vermicularis Enterocytozoon_bieneusi EpsteinBarr_virus Equus_caballus Eremothecium_cymbalariae Erinaceus_europaeus Erysiphe_necator Eucalyptus_grandis Eutrema_salsugineum Eutypa_lata Exophiala_aquamarina Exophiala_dermatitidis Exophiala_mesophila Exophiala_oligosperma Exophiala_sideris Exophiala_spinifera Exophiala_xenobiotica Fasciola_hepatica Felis_catus Fibroporia_radiculosa Ficedula_albicollis Fomitopsis_pinicola Fonsecaea_pedrosoi Fragaria_vesca Fragilariopsis_cylindrus Fusarium_fujikuroi Fusarium_graminearum Fusarium_oxysporum Fusarium_pseudograminearum Fusarium_solani Fusarium_verticillioides Gadus_morhua Gaeumannomyces_graminis Galerina_marginata Gallid_herpesvirus_2 Gallus_gallus Gasterosteus_aculeatus Geospiza_fortis Giardia_lamblia Glarea_lozoyensis Globodera_pallida Gloeophyllum_trabeum Glycine_max Gongylonema_pulchrum Gorilla_gorilla Gossypium_raimondii Grosmannia_clavigera Guillardia_theta Gymnopus_luxurians Haemonchus_contortus Haemonchus_placei Halocynthia_roretzi Halyomorpha_halys Harpegnathos_saltator Hebeloma_cylindrosporum Helianthus_annuus Heliconius_melpomene Heligmosomoides_bakeri Helobdella_robusta Hemiselmis_andersenii Heterobasidion_annosum Heterobasidion_irregulare Heterodontus_francisci Heterorhabditis_bacteriophora Histoplasma_capsulatum Homo_sapiens Hordeum_vulgare Human_herpesvirus_2 Hyaloperonospora_arabidopsidis Hydatigera_taeniaeformis Hydnomerulius_pinastri Hydra_magnipapillata Hymenolepis_diminuta Hymenolepis_microstoma Hymenolepis_nana Ictidomys_tridecemlineatus Ipomoea_batatas Ixodes_scapularis Jaapia_argillacea Kazachstania_africana Kazachstania_naganishii Kluyveromyces_delphensis Kluyveromyces_lactis Kluyveromyces_thermotolerans Kluyveromyces_waltii Komagataella_pastoris Kuraishia_capsulata Laccaria_amethystina Laccaria_bicolor Lachancea_kluyveri Lachancea_lanzarotensis Lachancea_thermotolerans Latimeria_chalumnae Leersia_perrieri Leishmania_braziliensis Leishmania_infantum Leishmania_major Leishmania_mexicana Lepisosteus_oculatus Leptinotarsa_decemlineata Leptosphaeria_maculans Lingula_unguis Linum_usitatissimum Litomosoides_sigmodontis Loa_loa Lodderomyces_elongisporus Lottia_gigantea Lotus_japonicus Loxodonta_africana Lucilia_cuprina Lutzomyia_longipalpis Lycopersicon_esculentum Macaca_fascicularis Macaca_mulatta Macrophomina_phaseolina Macropus_eugenii Magnaporthe_oryzae Magnaporthe_poae Malassezia_globosa Malassezia_sympodialis Malus_domestica Manihot_esculenta Marmota_monax Marssonina_brunnea Medicago_sativa Medicago_truncatula Megaselia_scalaris Melampsora_laricipopulina Meleagris_gallopavo Melitaea_cinxia Meloidogyne_floridensis Meloidogyne_hapla Meloidogyne_incognita Mesocestoides_corti Metarhizium_acridum Metarhizium_album Metarhizium_anisopliae Metarhizium_brunneum Metarhizium_guizhouense Metarhizium_majus Metarhizium_robertsii Meyerozyma_guilliermondii Microbotryum_violaceum Microcebus_murinus Micromonas_pusilla Micromonas_sp_RCC299 Microsporidia_sp Microsporum_canis Microsporum_gypseum Millerozyma_farinosa Mimulus_guttatus Mixia_osmundae Mnemiopsis_leidyi Moniliophthora_perniciosa Moniliophthora_roreri Monodelphis_domestica Monosiga_brevicollis Mucor_circinelloides Musa_acuminata Musca_domestica Mustela_putorius_furo Mus_musculus Myceliophthora_thermophila Mycosphaerella_fijiensis Mycosphaerella_graminicola Myotis_brandtii Myotis_lucifugus Naegleria_gruberi Nasonia_vitripennis Naumovozyma_castellii Naumovozyma_dairenensis Necator_americanus Nectria_haematococca Nematocida_parisii_ertm1 Nematocida_sp Nematostella_vectensis Neofusicoccum_parvum Neosartorya_fischeri Neospora_caninum Neurospora_crassa Neurospora_discreta Neurospora_tetrasperma Nicotiana_sp. Nicotiana_tabacum Nippostrongylus_brasiliensis Nomascus_leucogenys Nosema_apis Nosema_bombycis Nosema_ceranae Ochotona_princeps Octopus_bimaculoides Oesophagostomum_dentatum Ogataea_parapolymorpha Oidiodendron_maius Oikopleura_dioica Onchocerca_flexuosa Onchocerca_ochengi Onchocerca_volvulus Oncorhynchus_tshawytscha Ophiocordyceps_sinensis Ophiostoma_piceae_uamh Ophisaurus_gracilis Opisthorchis_viverrini Ordospora_colligata Oreochromis_niloticus Ornithorhynchus_anatinus Oryctolagus_cuniculus Oryza_barthii Oryza_brachyantha Oryza_glaberrima Oryza_glumaepatula Oryza_indica Oryza_longistaminata Oryza_meridionalis Oryza_nivara Oryza_punctata Oryza_rufipogon Oryza_sativa Oryzias_latipes Ostreococcus_lucimarinus Ostreococcus_RCC809 Ostreococcus_tauri Otolemur_garnettii Ovis_aries Oxytricha_trifallax Panicum_virgatum Pan_paniscus Pan_troglodytes Papio_anubis Paracoccidioides_brasiliensis Paracoccidioides_sp_lutzii Paramecium_tetraurelia Parascaris_equorum Parastrongyloides_trichosuri Patiria_miniata Paxillus_involutus Paxillus_rubicundulus PBM_CONSTRUCTS Pediculus_humanus Pelodiscus_sinensis Penicillium_chrysogenum Penicillium_digitatum Penicillium_expansum Penicillium_italicum Penicillium_marneffei Penicillium_oxalicum Penicillium_rubens Penicillium_solitum Pestalotiopsis_fici Petromyzon_marinus Petroselinum_crispum Petunia_x_hybrida Phaeodactylum_tricornutum Phaeosphaeria_nodorum Phanerochaete_carnosa Phanerochaete_chrysosporium Phaseolus_vulgaris Phlebiopsis_gigantea Phlebotomus_papatasi Phoenix_dactylifera Phycomyces_blakesleeanus Physcomitrella_patens Phytophthora_capsici Phytophthora_infestans Phytophthora_kernoviae Phytophthora_lateralis Phytophthora_parasitica Phytophthora_ramorum Phytophthora_sojae Pichia_angusta Pichia_kudriavzevii Pichia_pastoris Pichia_stipitis Piloderma_croceum Piriformospora_indica Pisolithus_microcarpus Pisolithus_tinctorius Pisum_sativum Plasmodium_berghei Plasmodium_chabaudi Plasmodium_falciparum Plasmodium_knowlesi Plasmodium_vivax Plasmodium_yoelii Pleurobrachia_pileus Pleurotus_djamor Pleurotus_ostreatus Plicaturopsis_crispa Pneumocystis_carinii Pneumocystis_murina Podocoryne_carnea Podospora_anserina Poecilia_formosa Pogona_vitticeps Polysphondylium_pallidum Pongo_abelii Populus_trichocarpa Postia_placenta Pristionchus_exspectatus Pristionchus_pacificus Procavia_capensis Protopolystoma_xenopodis Prunus_mume Prunus_persica Pseudocercospora_fijiensis Pseudogymnoascus_destructans Pseudogymnoascus_pannorum Pseudozyma_antarctica Pseudozyma_aphidis Pseudozyma_brasiliensis Pseudozyma_flocculosa Pseudozyma_hubeiensis Pteropus_vampyrus Puccinia_graminis Puccinia_triticina Punctularia_strigosozonata Pyrenophora_teres Pyrenophora_triticirepentis Pythium_aphanidermatum Pythium_arrhenomanes Pythium_irregulare Pythium_iwayamai Pythium_ultimum Pythium_vexans Rattus_norvegicus Rhabditophanes_kr3021 Rhinocladiella_mackenziei Rhizoctonia_solani Rhizophagus_irregularis Rhizopus_oryzae Rhodnius_prolixus Rhodosporidium_toruloides Rhodotorula_glutinis Ricinus_communis Romanomermis_culicivorax Rozella_allomycis Saccharomyces_arboricola Saccharomyces_bayanus Saccharomyces_castellii Saccharomyces_cerevisiae Saccharomyces_kudriavzevii Saccharomyces_mikatae Saccharomyces_paradoxus Saccharomycetaceae_sp_ashbya_aceri Saccoglossus_kowalevskii Salix_purpurea Salpingoeca_rosetta Saprolegnia_parasitica Sarcophilus_harrisii Sarsia_sp._Long_Island_Sound Scedosporium_apiospermum Scheffersomyces_stipitis Schistocephalus_solidus Schistosoma_curassoni Schistosoma_haematobium Schistosoma_japonicum Schistosoma_mansoni Schistosoma_margrebowiei Schistosoma_mattheei Schistosoma_rodhaini Schizophyllum_commune Schizosaccharomyces_cryophilus Schizosaccharomyces_japonicus Schizosaccharomyces_octosporus Schizosaccharomyces_pombe Schmidtea_mediterranea Scleroderma_citrinum Sclerotinia_borealis Sclerotinia_sclerotiorum Selaginella_moellendorffii Serendipita_vermifera Serpula_lacrymans Setaria_italica Setosphaeria_turcica Soboliphyme_baturini Solanum_lycopersicum Solanum_tuberosum Solenopsis_invicta Sordaria_macrospora Sorex_araneus Sorghum_bicolor Spadella_cephaloptera Spathaspora_passalidarum Sphaerobolus_stellatus Sphaerulina_musiva Spirometra_erinaceieuropaei Spizellomyces_punctatus Sporisorium_reilianum Sporobolomyces_roseus Sporothrix_brasiliensis Sporothrix_schenckii Spraguea_lophii Stachybotrys_chartarum Stachybotrys_chlorohalonata Stagonospora_nodorum Steinernema_carpocapsae Steinernema_feltiae Steinernema_glaseri Steinernema_monticolum Steinernema_scapterisci Strigamia_maritima Strongylocentrotus_purpuratus Strongyloides_papillosus Strongyloides_stercoralis Strongyloides_venezuelensis Strongylus_vulgaris Suillus_luteus Sus_scrofa Syphacia_muris Taenia_asiatica Taenia_solium Taeniopygia_guttata Takifugu_rubripes Talaromyces_marneffei Talaromyces_stipitatus Tarsius_syrichta Teladorsagia_circumcincta Tetrahymena_thermophila Tetranychus_urticae Tetraodon_nigroviridis Tetrapisispora_blattae Tetrapisispora_phaffii Thalassiosira_pseudonana Thecamonas_trahens Theileria_annulata Theileria_parva Thelazia_callipaeda Theobroma_cacao Thielavia_terrestris Tilletiaria_anomala Togninia_minima Torrubiella_hemipterigena Torulaspora_delbrueckii Toxocara_canis Toxoplasma_gondii Trachipleistophora_hominis Trametes_cinnabarina Tremella_mesenterica Tribolium_castaneum Trichinella_nativa Trichinella_spiralis Trichobilharzia_regenti Trichoderma_atroviride Trichoderma_reesei Trichoderma_virens Trichomonas_vaginalis Trichophyton_equinum Trichophyton_interdigitale Trichophyton_rubrum Trichophyton_soudanense Trichophyton_tonsurans Trichophyton_verrucosum Trichoplax_adhaerens Trichosporon_asahii Trichuris_muris Trichuris_suis Trichuris_trichiura Trionyx_sinensis Tripedalia_cystophora Triticum_aestivum Triticum_urartu Trypanosoma_brucei Trypanosoma_congolense Trypanosoma_cruzi Trypanosoma_vivax Tuber_melanosporum Tulasnella_calospora Tupaia_belangeri Tursiops_truncatus Tyto_alba Uncinocarpus_reesii Ustilaginoidea_virens Ustilago_hordei Ustilago_maydis Vanderwaltozyma_polyspora Vavraia_culicis Verruconis_gallopava Verticillium_albo_atrum Verticillium_alfalfae Verticillium_dahliae Vicugna_pacos Vitis_vinifera Vittaforma_corneae Volvox_carteri Wallemia_ichthyophaga Wallemia_sebi Wickerhamomyces_ciferrii Wuchereria_bancrofti Xenopus_laevis Xenopus_tropicalis Xiphophorus_maculatus Yarrowia_lipolytica Zea_mays Zootermopsis_nevadensis Zygosaccharomyces_bailii Zygosaccharomyces_rouxii Zymoseptoria_tritici
By Domain Type	ABF1 AFT AP-2 AP2 APSES ARID/BRIGHT AT hook B3 BBR BED ZF bHLH Brinker bZIP C2H2 ZF C2HC ZF CBF/NF-Y CCCH ZF CENPB CG-1 Copper fist CSD CSL CUT CxxC DM Dof E2F EBF1 EIN3 Ets FAR1 FLYWCH Forkhead GATA GCM GCR1 Grainyhead GRAS GTF2I-like Homeodomain HSF IBD ICP4 IRF Leafy LOB MADF MADS box MATalpha_HMGbox MBD mTERF Myb NAC/NAM Ndt80/PhoG NFX Nuclear receptor p53 Paired box Pipsqueak PLATZ POU Prospero Rap1 Rel RFX RRM Runt RWP-RK S1FA SAND SBP Sigma 70 SMAD Sox STAT STE Storekeeper T-box TBP TCP TCR/CxC TEA THAP finger Unknown VHR1 Whirly WRC WRKY Zinc cluster
By Motif Evidence	Direct Direct & Inferred--> Inferred None
By Evidence Type	B1H ChIP-chip ChIP-exo ChIP-seq ChIP-seq+ChIP-exo COMPILED Dap-seq DNaseI footprinting EMSA Misc PBM PBM, CSA and or DIP-chip SELEX SMiLE-seq Transfac Unknown
By Study	Badis,2008 Badis,2009 Barazandeh,2018 Barrera,2016 Berger,2006 Berger,2008 Busser,2012b Busser,2012a Campbell,2010 Chang,2013 Cheatle-Jarvela,2014 Chen,2011 Contrino,2012 DeBoer,2011 DelBianco,2010 DeMasi,2011 DeSilva,2008 Franco-Zorrilla,2014 Gabut,2011 Gerstein,2012 Gordan,2011 Grove,2009 Guo,2014 Hallikas,2006 Heinz,2010 Isakova,2017 Jolma,2010 Jolma,2013 Kulakovskiy,2009 Kulakovskiy,2013 Lam,2011 Lambert,2019 LehtiShiu,2015 Lesch,2009 Li,2017 Lindemose,2014 Liu,2018 Mann,2013 Mathelier,2014 Matys,2006 Ming,2013 Najafabadi,2015b Najafabadi,2015a Nakagawa,2013 Narasimhan,2015 Nitta,2015 OMalley,2016 Peterson,2012 Radke,2013 Rhee,2011 Rodriguez-Martinez,2017 Scharer,2009 Schmitges,2016 SebePedros,2013 Siggers,2014 Soruco,2013 Sullivan,2014 Tillo,2017 Wei,2010 Weirauch,2014 Weirauch,2013 Yin,2017b Yin,2017 Zhu,2011 Zhu,2009
Database Build	Version 2.00 Version 1.02 Version 1.01 Version 0.90
Database build 2.00 now available!

Last updated: Jan 8th, 2019 Database Build 2.00

Current content: 11491 motifs. 165030 TFs with at least one binding motif (4559 from direct experiments), out of a total of 392333 TFs from 321 families in 741 species